A public filing by AquaBounty has revealed that Edinburgh-based tycoon Yuriy Lopatynskyy has invested in the genetically modified (GM) salmon producer.

Luxembourg-based LLF Financial, half-owned by Lopatynskyy, purchased just under 3.2 million shares via two companies -- Sopica Special Opportunities and Sopica Global Retail Growth Fund -- giving the group 9.9 percent of the outstanding shares.

The other half of LLF Financial is owned by Ukraine-based Hennadiy Lopatynskyy. It's unclear the relation between the two men.

The UK-based Lopatynskyy is linked to several investments and properties around the globe, including multiple offshore groups cited in the Paradise Papers, among them Uraslib Russia Select Fund, USB Asset Management, Sopica CIS Small Cap Fund, FMIG and BAMI First Mercantile.

Lopatynskyy also owns Northern Link, which was the largest shareholder of the Scottish Salmon Company. Northern Link is a wholly-owned subsidiary of the Virgin Islands-based Sopica Global Retail Growth Fund, which is advised by LLF Financial.

Lopatynskyy took a controlling interest in Scottish salmon farmer Lighthouse Caledonia in 2009 through his private equity firm Northern Link, right before it was acquired by SSC.

Northern Link has held in the past several other investments in aquaculture and marine-related companies in Norway, Peru and Russia, including stakes in Leroy Seafood, Codfarmers, Marine Farms, Copeinca and Villa Organic.